Revised submission

This release represents all the edits made to the manuscript to address reviewer comments from our initial submission to PLoS Comp Bio.

Initial submission to plos comp bio

This version of the manuscript has been submitted to PLoS Comp Biology. The tracking number for this submission is: PCOMPBIOL-D-16-00565

PeerJ pre-print (first version)

Version of the manuscript submitted on October 23rd, 2015 as a pre-print to PeerJ